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Fig. 2. Effect of position along the hydrologic gradient and topsoil status on (a) soil moisture, (b) bulk density, (c) soil organic matter, (d) clay, (e) silt, (f) sand, (h) water holding capacity, (h) the P sorption index, and (i) microbial biomass C. Data presented are means and error bars represent ±1 standard error. Data points with different letters are significantly different at the p < 0.10 level according to a least-squared differences test.