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a Bioforsk, Norwegian Institute for Agricultural and Environmental Research, Soil and Environment Division, 1432 Ås, Norway
b UMR Biogéochimie et Ecologie des Milieux Continentaux. (BioEMCo), INRA-CNRS-INAPG-ENS-ENSCP-Univ. Paris VI, Bâtiment EGER, 78850 Thiverval-Grignon France
c Norwegian Univ. of Life Sciences, Dep. of Plant and Environmental Sciences, 1432 Ås, Norway
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