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a Ecole Supérieure d'Agriculture de Purpan75, voie du TOEC BP 57 611, 31 076 Toulouse Cedex 3, France
b UMR INRA/INAPG Environment and Arable Crops, Institut National de la Recherche Agronomique/Institut National Agronomique Paris-Grignon, BP 01, 78 850 Thiverval-Grignon, France
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